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A procession of segmented creatures moves across the sea floor.
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Up ahead, hundreds have begun shedding their exoskeletons
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and piling on top of one another in what appears to be... a massive orgy.
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But this is not some alien world.
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Itโs Earth about 500 million years ago,
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when these creatures, called trilobites, prospered.
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Prevailing for around 270 million years
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and encompassing more than 20,000 distinct species,
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trilobites are some of the most successful lifeforms in Earthโs history.
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When they sprung into existence,
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they were among the most diverse and sophisticated organisms
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Earth had yet seen.
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And, as the earliest known animals with complex eyes,
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trilobites had a unique perspective on the ancient world.
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For almost all of Earthโs history before the rise of the trilobites,
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life had mostly consisted of microscopic marine organisms.
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But then, scientists think an increase in oxygen allowed multicellular lifeforms
1:17
to extract more energy from their food and perform more complex functions.
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This then enabled the rise of carnivores,
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which in turn spurred a productive arms race,
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resulting in whatโs known as the Cambrian explosion.
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Within about 20 million years,
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life had branched out to include most of the animal groups we know today.
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Trilobites were an important part of this surge.
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All trilobites had three lengthwise lobes,
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but building off the same basic components,
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they varied greatly.
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Species ranged in length from a few millimeters to almost one meter
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and some were equipped with intriguing ornaments.
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As a result, they filled many distinct niches over the ages.
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Many trilobites plowed or burrowed into the sea floor while others swam freely.
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Certain species had spines, horns, and even protruding tridents.
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And their social behavior was complex:
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they came together to search for food, find safety in numbers, migrate and mate.
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In fact, their fossilized conga lines represent
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some of the first evidence of animal group behaviour.
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Like modern arthropods,
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trilobites had compound eyes composed of many tiny lenses.
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But theirs were made of the mineral calcite,
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which also constituted their exoskeletons.
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These lenses allowed trilobites to form sharp images
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and quickly sense changes in light.
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Some had long eye stalks that helped them peep above the muddy sea floor
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as they burrowed beneath.
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Others had large dragonfly-like eyes
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that they probably used to scour the ocean bottom
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as they swam upside-down in dim waters.
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And some trilobites had pillar-like eyes that may have offered 360 degree views,
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along with structures that provided shade from overhead light.
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Yet despite their dominance of the seas,
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trilobites were vulnerable to environmental changes and predators.
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About 444 million years ago, Earth cooled and sea levels dropped,
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radically changing some of their habitats.
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This was the beginning of what would prove to be the trilobiteโs long demise.
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Some 20 million years later, fish with jaws began sweeping the sea.
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Trilobites developed spiky ornamentation
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and neat ways of locking their joints shut.
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Many species could curl themselves into balls,
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entirely sealing their soft parts inside their hard exoskeletons.
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Then, another extinction event shook the world.
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Trilobite diversity dwindled, and come about 360 million years ago,
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only one of the 10 former trilobite orders remained.
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Finally, rapid climate change spurred
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the greatest known mass extinction event in Earth's history.
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This catastrophic period finished the trilobites offโ
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along with approximately 96% of all marine species.
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But trilobites left a remarkable record behind.
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Their calcite exoskeletons made for hardy fossils
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that would remain intact for hundreds of millions of years to come.
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Weโve found trilobite fossils on every single continentโ
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many in unlikely environments that were once part of the ocean floor.
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Trilobites have no direct descendants,
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but their evolutionary cousins are alive and well.
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And since arthropods make up over 80% of current animal species,
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we might say that, although Earth may no longer be the planet of the trilobites,
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their distant relatives still reign supreme.